SAVANNAH — Authorities in Savannah say a truck driver has been cited after a trailer containing eight circus animals became unhitched on a highway.

Authorities said late Monday that Michael K. Ramos of Riverview, Fla., was charged with driving too fast for conditions, failing to maintain his lane and having an improper class license after the accident on Interstate 516.

Police say Ramos hit a sign, which dislodged the trailer containing animals performing in the UniverSoul Circus. Police said medics treated two llamas for minor nose and foot injuries.

The eastbound lanes of the highway were closed for more than five hours after the 4 p.m. accident between the exit and entrance ramps to Veterans Parkway.

Firefighters helped transfer the llamas, two zebras, two ponies and two camels to another trailer.